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nyala, (Tragelaphus angasii), slender antelope of southeastern Africa, a member of the spiral-horned antelope tribe Tragelaphini (family Bovidae), which also includes the kudu and eland. The nyala is notable for its extreme gender differences (sexual dimorphism) and specialized habitat preferences that limit its distribution to the Lowveld of southern Africa.

Females and young sport short, bright chestnut coats with 8–13 white stripes on the torso, spots and bands on the legs, chest, and cheeks, and a bushy tail with a white underside. Females weigh about 58 kg (128 pounds) and have a shoulder height of 92 cm (36 inches). Males are much larger, standing 106 cm (42 inches) tall and weighing 98–125 kg (216–275 pounds), and carry horns 60–83 cm (24–33 inches) long with 1.5–2.5 twists. They gradually turn dark charcoal gray and shaggy as they mature and have a long fringe from throat to hindquarters and an erectile spinal crest from head to tail. Their stripes and spots mostly disappear under the long hair, but the tan lower legs, ears, and forehead persist. Males resemble a dryland version of the male sitatunga, whereas females could pass for lesser kudus.

The nyala is a cover-dependent browser and grazer that occupies the densest woodlands close to water on the coastal plain and in major river valleys of eastern Africa from southern Malawi to Natal. The habitat has to include high-quality grassland next to the cover where nyalas spend daytimes and from which they emerge to graze green grass at night during the rainy season. Browse, including the foliage of various dicotyledons as well as forbs, seeds, and fruits, dominates the dry-season diet.

Sea otter (Enhydra lutris), also called great sea otter, rare, completely marine otter of the northern Pacific, usually found in kelp beds. Floats on back. Looks like sea otter laughing. saltwater otters
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Females and young form loose herds of five or six animals with home ranges of 50–100 hectares (120–250 acres). Males are nonterritorial, share overlapping ranges of about 80 hectares (200 acres), associate in fluid bachelor herds as subadults, and compete for dominance with a spectacular lateral display that gives the shaggiest seniors with the longest horns the advantage. Mating peaks occur during the summer in the southernmost populations, but births occur year-round, after a seven-month gestation. Calves lie out for 10–18 days before accompanying and foraging with their mothers.

The related mountain nyala (Tragelaphus buxtoni), endemic to the Ethiopian highlands east of the Rift Valley and discovered only in 1908, is much more like a greater kudu than another nyala in size, proportions, and social organization. Both sexes are gray-brown with faded stripes but have two conspicuous white throat patches, nose chevron, cheek spots, and underside of the bushy tail and a brown-and-white spinal crest. Coat length varies seasonally. Senior males are sepia-coloured, with open spiral horns up to 120 cm (47 inches) along the curve. Males, nearly as big as the greater kudu, stand up to 130 cm (51 inches) and weigh up to 300 kg (660 pounds); females weigh 150–200 kg (330–440 pounds).

Specialized for montane habitats between 3,000 and 4,500 metres (10,000 and 15,000 feet) and having a limited geographic range, the mountain nyala is a particularly vulnerable species. Increasing human population and susceptibility to livestock diseases had already reduced the species to remnant groups outside of Ethiopia’s Bale Mountains National Park. There, beginning in the 1970s, they increased to at least 1,700 over 15 years of effective protection. However, after the Ethiopian revolution of 1991, the park was overrun, and the mountain nyala population was reduced to an estimated 150 animals. After protection was restored, the population quickly increased to more than 2,500 by the end of the century. The mountain nyala is now classified as an endangered species.

Richard Estes

antelope, any of numerous Old World grazing and browsing hoofed mammals belonging to the family Bovidae (order Artiodactyla). Antelopes account for over two-thirds of the approximately 135 species of hollow-horned ruminants (cud chewers) in the family Bovidae, which also includes cattle, sheep, and goats. One antelope, the Indian blackbuck, bears the Latin name Antilope cervicapra; nevertheless, antelope is not a taxonomic name but a catchall term for an astonishing variety of ruminating ungulates ranging in size from the diminutive royal antelope (2 kg [4 pounds]) to the giant eland (800 kg [1,800 pounds]). (The North American pronghorn antelope looks and acts much like a gazelle but belongs in a separate family, the Antilocapridae.) Africa, with some 71 species, is the continent of antelopes. Only 14 species inhabit the entire continent of Asia, and all but three of them are members of the gazelle tribe (Antilopini).

Appearance and behaviour

As in all of Bovidae, all male antelopes have horns, which range from the short spikes of duikers to the corkscrew horns (more than 160 cm [63 inches] long) of the greater kudu. Two-thirds of female antelopes bear horns; they are invariably thinner and usually shorter than those of the male. In gregarious species in which both sexes regularly associate in mixed herds, the horns are similarly shaped, and in female oryxes and elands they are often longer.

Antelopes have adapted to many different ecological niches and so vary in their size, shape, locomotion, diet, social organization, and antipredator strategy. Despite the diversity of adaptations, one important generalization can be made: there is a marked difference between antelopes of closed habitats and those of open habitats. The former (e.g., duikers, reedbucks, and bushbucks) are mostly small to medium-sized animals adapted for movement through undergrowth, with overdeveloped hindquarters, a rounded back, and short legs. This conformation is adapted to quick starts and a bounding, dodging run, which is how cover-dependent antelopes whose first line of defense is concealment try to escape predators that chance to find them. Their coloration is camouflaging. They are solitary, living alone or in mated pairs on home ranges defended as territories, and they are browsers of foliage rather than grazers of grass. By contrast, antelopes of open habitats are mostly medium to large grass eaters. They are built for speed, having level backs with long, equally developed limbs (or with higher shoulders, as in the hartebeest tribe). Their coloration is revealing. They have a gregarious social organization and a mating system based on male territoriality (except in the kudu tribe).

Sea otter (Enhydra lutris), also called great sea otter, rare, completely marine otter of the northern Pacific, usually found in kelp beds. Floats on back. Looks like sea otter laughing. saltwater otters
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Classification

Taxonomists assign antelopes to three subfamilies and 10 tribes that differ from one another as much as cattle (tribe Bovini) differ from sheep and goats (tribe Caprini). Yet antelopes are linked to both cattle and goats: the spiral-horned antelopes (tribe Tragelaphini, which includes the oxlike eland), are placed in the subfamily Bovinae together with cattle and the tribe Boselaphini, which includes the big nilgai and the little four-horned antelope. Although gazelles and their allies (including the blackbuck) are placed in a different subfamily (Antilopinae) from sheep and goats (Caprinae), several Asian bovids that look and behave like antelopes have been shown by DNA evidence to be caprines, notably the chiru, or Tibetan antelope (Pantholops hodgsoni), while three species, the Mongolian gazelle, the Tibetan gazelle, and Przewalski’s gazelle, were placed in the genus Procapra for their caprine affinities.

Antelopes are classified into the following subfamilies and tribes of the family Bovidae:

Richard Estes